Abstract:Recursive processing in sentence comprehension is considered a hallmark of human linguistic abilities. However, its underlying neural mechanisms remain largely unknown. We studied whether a recurrent neural network with Long Short-Term Memory units can mimic a central aspect of human sentence processing, namely the handling of long-distance agreement dependencies. Although the network was solely trained to predict the next word in a large corpus, analysis showed the emergence of a small set of specialized units that successfully handled local and long-distance syntactic agreement for grammatical number. However, simulations showed that this mechanism does not support full recursion and fails with some long-range embedded dependencies. We tested the model's predictions in a behavioral experiment where humans detected violations in number agreement in sentences with systematic variations in the singular/plural status of multiple nouns, with or without embedding. Human and model error patterns were remarkably similar, showing that the model echoes various effects observed in human data. However, a key difference was that, with embedded long-range dependencies, humans remained above chance level, while the model's systematic errors brought it below chance. Overall, our study shows that exploring the ways in which modern artificial neural networks process sentences leads to precise and testable hypotheses about human linguistic performance.