We describe Bayesian Layers, a module designed for fast experimentation with neural network uncertainty. It extends neural network libraries with layers capturing uncertainty over weights (Bayesian neural nets), pre-activation units (dropout), activations ("stochastic output layers"), and the function itself (Gaussian processes). With reversible layers, one can also propagate uncertainty from input to output such as for flow-based distributions and constant-memory backpropagation. Bayesian Layers are a drop-in replacement for other layers, maintaining core features that one typically desires for experimentation. As demonstration, we fit a 10-billion parameter "Bayesian Transformer" on 512 TPUv2 cores, which replaces attention layers with their Bayesian counterpart.